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The Order of the Guardians is a group of Trills who care for the Trill symbionts in the Caves of Mak'ala. They rarely leave the Caves and have a reputation of being odd. (DS9 novel: Trill: Unjoined)

In 2371, Jadzia Dax, Benjamin Sisko and Julian Bashir consulted the Guardian Timor for information on hallucinations Jadzia had been having. (DS9 episode: "Equilibrium")

The Guardians preside over the zhian'tara ceremony, transferring the memories and personality of past hosts into the bodies of participants. (DS9 episode: "Facets")

Ranul Keru was once a member. (DS9 novel: Trill: Unjoined)

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